A−1 is characterized by
AA−1=I
Using the interpretation from part (a), we can interpret 
A−1 as follows: it gives the required herbivores-eaten-by-carnivores coefficients so that if 
B=A−1 then each plant 
type gets eventually consumed exactly once by a 
unique carnivore.
Likewise, 
B−1 gives the required plants-eaten-by-herbivores coefficients so that if 
A=B−1 then each plant 
type gets eventually consumed exactly once by a 
unique carnivore.
Unclear to me the significance of 
(AB)−1